The problem was that the PA announcer and most of the fans thought that the runner/receiver had stretched for a first down before going out of bounds on the previous second-down play. It seems like UConn thought the same thing. Instead of running a short yardage play to gain a half-yard or so, they hurried into that third-down pass play 10 yards downfield, which almost turned into an INT.
However, the refs had never moved the chains downfield, because they'd ruled that the runner on the second down play had gone down short of the line to gain. They had it as being third down on that near-INT pass play, not first down. So, the incomplete pass made it 4th and 1, and UConn chose to punt.
It sucked. I thought the refs should have stopped play when they heard the PA announcer call "first down" to the entire stadium to clarify the spot, and that the runner hadn't gotten a first down. I can't recall ever seeing anything like that happen before, to be honest, and I've been watching football since Jimmy Carter was president. UConn should have realized that the chains hadn't moved and backed off from the pass play they'd called, but the refs really could have averted the confusion as well.